I saw a documentary about birds...I have a dvd of it somewhere, I think...they used a camera on a light flying craft to film geese in flight, and you could see the geese get bounced out of formation if they drifted out of the flow. There was a turbulence, I forget how they illustrated it, and as long as the geese stayed in that level V, they were almost pulled along in a slipstream. But as soon as the lead goose got tired, it would drift out of point, get buffeted around a bit, then slip back into place behind the new leader. There was a brief wobble as the V reformed for all the geese in the formation. If a goose got more out of place, it was almost ejected from the V. It was fascinating.